Advancing knowledge. Supporting conservation.


Requests for reprints: 


  • Bulla, M., J. Reneerkens, E.L. Weiser, A. Sokolov, A.R. Taylor, B. Sittler, B.J. McCaffery, D.R. Ruthrauff, D.H. Catlin, D.C. Payer, et al. 2019. Comment on “Global pattern of nest predation is disrupted by climate change in shorebirds.” Science 364(6445):eaaw8529. DOI:10.1126/science.aaw8529

  • Zeigler, S.L., B.T. Gutierrez, E.J. Sturdivant, D.H. Catlin, J.D. Fraser, A. Hecht, S.M. Karpanty, N.G. Plant, E.R. Thieler. 2019. Using a Bayesian network to understand the importance of coastal storms and undeveloped landscapes for the creation and maintenance of early successional habitat. PLOS ONE 14(7):e0209986. DOI:10.1371/journal.pone.0209986

  • Davis K.*, S.M. Karpanty, J. Spendelow, J. Cohen, M. Althouse, K. Parsons, C. Luttazi, D.H. Catlin, and D. Gibson. 2019. Residency, recruitment, and stopover duration of hatch-year Roseate Terns (Sterna dougallii) during the pre-migratory staging period. Avian Conservation and Ecology: 14 (2):11. DOI:10.5751/ACE-01416-140211.

  • Weithman, C.E.*, S.G. Robinson*, K.L. Hunt, J. Altman, H.A. Bellman, A.L. DeRose-Wilson, K.M. Walker*, J.D. Fraser, S.M. Karpanty, and D.H. Catlin. 2019. Growth of two Atlantic Coast Piping Plover populations. The Condor: Ornithological Applications. Early View.

  • Robinson, S.G.*, H.A. Bellman, K. Thyberg, J.D. Fraser, D.H. Catlin, and S.M. Karpanty. 2019. Observations of double brooding and inbreeding of piping plovers on Fire and Westhampton Islands, New York. Northeastern Naturalist 26 (3): N38–N42.

  • Robinson, S.G.*, J.D. Fraser, D.H. Catlin, S. Karpanty, J. Altman, R. Boettcher, K. Holcomb, C. Huber, K.L. Hunt, and A. Wilke. 2019. Irruptions: evidence for breeding season habitat limitation in Piping Plover (Charadrius melodus). Avian Conservation and Ecology 14 (1):19. DOI:10.5751/ACE-01373-140119

  • Gibson, D., A.D. Hornsby, M.B. Brown, J. Cohen, L. Dinan, J.D. Fraser, M. Friedrich, C.L. Gratto-Trevor, K.L Hunt, M. Jeffery, J. Jorgensen, P. Paton, S. Robinson, J. Rock, M. Stantial, C. Weithman, and D.H. Catlin. 2019. Migratory bird adheres to Bergmann’s Rule by responding to environmental conditions throughout the annual lifecycle. Ecography 42(8): 1482–1493. DOI:10.1111/ecog.04325

  • Walker, K.M.*, J.D. Fraser, D.H. Catlin, S.J. Ritter, S.G., Robinson*, H.A. Bellman, A. DeRose-Wilson, S.M. Karpanty, and S.T. Papa. 2019. Hurricane Sandy and engineered response created habitat for a threatened shorebird. Ecosphere:10(6):e02771. DOI:10.1002/ecs2.2771

  • Catlin, D.H., D. Gibson, K.L. Hunt, M.J. Friedrich, C.E. Weithman*, S.M. Karpanty, and J.D. Fraser. 2019. Direct and indirect effects of nesting density on survival and breeding propensity of an endangered shorebird. Ecosphere: 10(6):e02740. DOI: 10.1002/ecs2.2740.

  • Hunt, K. L., D. Gibson, M.J. Friedrich, C.J. Huber, J.D. Fraser, S.M. Karpanty, and D.H. Catlin. 2019. Using nest captures and video cameras to estimate survival and abundance of breeding piping plovers Charadrius melodus. Ibis: in press. DOI: 10.1111/ibi.12726

  • Fraser, J.D. and D.H. Catlin. 2019. Habitat Ecology and Conservation of Charadrius Plovers. Pp. 217–243 in Colwell and Haig (editors). The Population Ecology and Conservation of Charadrius Plovers (no. 52), CRC Press, Boca Raton, FL.

  • Heller, E.L.*, S.M. Karpanty, and J. D. Fraser. 2019. Depredation of a Snowy Egret Egretta thula by a Pomarine Jaeger Stercorarius pomarinus in Virginia. Marine Ornithology 47:29-31.    


  • Kwon, E., L. Houghton, R. Settlage, D.H. Catlin, S.M. Karpanty, and J.D. Fraser. 2018. Estimating transient populations of unmarked individuals at a migratory stopover site using generalized N-mixture models. Journal of Applied Ecology, 55(6): 2917–2932. DOI: 10.1111/1365-2664.13243.

  • DeRose-Wilson, A.L., K.L. Hunt, J.D. Monk**, D.H. Catlin, S.M. Karpanty, and J.D. Fraser. 2018. Piping plover chick survival negatively correlated with beach recreation. Journal of Wildlife Management, 82(8):1608–1616. DOI:10.1002./jwmg.21552.

  • Gibson, D. T.V. Riecke, T. Keyes, C. Depkin, J.D. Fraser, and D.H. Catlin. 2018. Application of Bayesian robust design model to assess the impacts of a hurricane on shorebird demography. Ecosphere 9(8):e02334. DOI: 10.1002/ecs2.2334.

  • Gibson, D., M.K. Chaplin, K.L. Hunt, M.J. Friedrich*, C.E. Weithman*, L.M. Addison, V. Cavalieri, S. Coleman, F.J. Cuthbert, J.D. Fraser, W. Golder, D. Hoffman, S.M. Karpanty, and D.H. Catlin. 2018. Impacts of anthropogenic disturbance on body condition, survival, and site fidelity of nonbreeding piping plovers. Condor 120: 566–580. DOI: 10.1650/CONDOR-17-148.1.

  • Nefas, S.M.**, K.L. Hunt*, J.D. Fraser, S.M. Karpanty, and D.H. Catlin. 2018. Least tern (Sternula antillarum) nest success and chick survival on the Missouri River following historic flooding. Wilson Journal of Ornithology, 130(2): 371–376. doi: 10.1676/16-210.1 

  • Kwon, E., J.D. Fraser, D.H. Catlin, S.M. Karpanty, C.E. Weithman*, and B. Muiznieks. 2018. Presence of ghost crabs and piping plover nesting success. Journal of Wildlife Management 82(4): 850–856. doi:10.1002/jwmg.21422.

  • Cohen, J.B., S.B. Maddock, M.K. Bimbi, W.W. Golder, O.E. LeDee, F.J. Cuthbert, D.H. Catlin, J.D. Fraser, and C.L. Gratto-Trevor. 2018. State uncertainty models and mark-resight models for understanding nonbreeding site use by piping plovers. Ibis 160: 342–354. doi:10.1111/ibi.12546.

  • Weithman, C.E.*, D. Gibson, K.M. Walker*, S. B. Maddock, J.D. Fraser, S.M. Karpanty, and D.H. Catlin. 2018. Discovery of an important stopover location for mirgratory piping plovers (Charadrius melodus) on South Point, Ocracoke Island, North Carolina. Waterbirds 41: 56–62.

  • Hunt, K.L.*, J.D. Fraser, M.J. Friedrich*, S.M. Karpanty, and D.H. Catlin. 2018. Demographic response of an imperiled bird suggests that engineered habitat restoration is no match for natural riverine processes. Condor: Ornithological Applications 120(1): 149–165. doi: 10.1650/CONDOR-17-93.1.


  • Wang, W., J.D. Fraser, and J. Chen. 2017. Distribution and long-term population trends of wintering waterbirds in Poyang Lake, China. Wetlands. doi:10.1007/s13157-017-0981-6.

  • Hunt, K.L.*, J.D. Fraser, S.M. Karpanty, and D.H. Catlin. 2017. Piping Plover (Charadrius melodus) body condition and prey abundance on flood-created habitat on the Missouri River, USA. Wilson Journal of Ornithology 129 (4): 754–764. doi: 10.1676/16-180.1.

  • Weithman, C.E.*, D. Gibson, K. Hunt*, M. Friedrich*, J. Fraser, S.M. Karpanty, and D.H. Catlin. 2017. Senescence and carryover effects of reproductive performance influence migration, condition, and breeding propensity. Ecology and Evolution 7: 11044–11056. doi:10.1002/ece3.3533.

  • Zeigler, S.L., E.R. Thieler, B.T. Gutierrez, N.G. Plant, M. Hines, J.D. Fraser, D.H. Catlin, S.M. Karpanty. 2017. Smartphone technologies and Bayesian networks to assess shorebird habitat selection. Wildlife Society Bulletin 41: 666–677. doi:10.1002/wsb.820.

  • Gibson, D., D.H. Catlin, K.L. Hunt, J.D. Fraser, S.M. Karpanty, M.J. Friedrich, M.K. Bimbi, J.B. Cohen, S.B. Maddock. 2017. Evaluating the impact of man-made disasters on imperiled species: piping plover and the Deepwater Horizon oil spill. Biological Conservation 212: 48–62.

  • Zeigler, S.L., D.H. Catlin, M. Bomberger Brown, J.D. Fraser, L.R. Dinan, K.L. Hunt, J.G. Jorgensen, and S.M. Karpanty. 2017. Effects of climate change and anthropogenic modification on a disturbance-dependent species in a large riverine system. Ecosphere 8(1):e01653. 10.1002/ecs2.1653

  • Davis, K.L.*, K.L. Schoenemann*, D.H. Catlin, K.L. Hunt, M.J. Friedrich, S.J. Ritter, J.D. Fraser, and S.M. Karpanty. 2017. Hatch-year piping plover (Charadrius melodus) prospecting and habitat quality influence second-year nest site selection. The Auk 134: 92–103.


  • Monk, J.D., A. DeRose-Wilson, J.D. Fraser, D.H. Catlin, and S.M. Karpanty. 2016. Observations of fish consumption by piping plovers. Northeastern Naturalist 23(3): 22–25.

  • Wang, W., J.D. Fraser, and J. Chen. 2016. Wintering waterbirds in the middle and lower Yangtze River floodplain: changes in abundance and distribution. Bird Conservation International 27: 167–186.

  • Catlin, D.H., S. Zeigler, M. Bomberger Brown, L.R. Dinan, J.D. Fraser, K.L. Hunt*, and J.G. Jorgensen. 2016. Metapopulation viability of an endangered shorebird depends on dispersal and human-created habitats: piping plovers (Charadrius melodus) and prairie rivers. Movement Ecology 4:6. 


  • DeRose-Wilson, A., Fraser, J. D., Karpanty, S. M. and Hillman, M. D. 2015. Effects of overflights on incubating Wilson's plover behavior and heart rate. The Journal of Wildlife Management 79(8): 1246–1254. 

  • Hunt, K.L.*, L.R. Dinan, M.J. Friedrich, M. Bomberger Brown, J.G. Jorgensen, D.H. Catlin, and J.D. Fraser. 2015. Density dependent double brooding in Great Plains piping plovers (Charadrius melodus). Waterbirds 38(4): 321–329.

  • Catlin, D.H., J.D. Fraser, and J.H. Felio*. 2015. Demographic responses of piping plovers to habitat creation on the Missouri River. Wildlife Monographs 192: 1–42. 

  • Friedrich, M.J.*, K.L. Hunt*, D.H. Catlin, and J.D. Fraser. 2015. The importance of site to mate choice: mate and site fidelity in piping plovers. Auk 132: 265–276. 


  • Catlin, D.H., O. Milenkaya, K.L. Hunt*, M.J. Friedrich*, and J. D. Fraser. 2014. Can river management improve the piping plover’s long-term survival on the Missouri River? Biological Conservation 180: 196–205.

  • McGowan, C.P., D. H. Catlin, T.L. Shaffer, C.L. Gratto-Trevor, and C. Aron. 2014. Establishing endangered species recovery criteria using predictive simulation modeling. Biological Conservation 177: 220–229.

  • Gieder, K.D.*, S.M. Karpanty, J.D. Fraser, D.H. Catlin, B.T. Gutierrez, N.G. Plant, A.M. Turecek, E.R. Thieler. 2014. A Bayesian network approach to predicting nest present of the federally-threatened piping plover (Charadrius melodus) using barrier island features.  Ecological Modelling: 276: 36–50. 


  • DeRose-Wilson, A.L.*, J.D. Fraser, S.M. Karpanty, and D.H. Catlin. 2013. Nest-site selection and demography of Wilson’s Plovers on a North Carolina barrier island. Journal of Field Ornithology 84(4): 329–344.

  • Fraser, J.D., S.M. Karpanty, J.B. Cohen, B.R. Truitt. 2013. The Red Knot (Calidris canutus rufa) decline in the Western Hempisphere: is there a lemming connection? Canadian Journal of Zoology 91: 13-16.

  • Hillman, M.D.*, S.M. Karpanty, J.D. Fraser, J.D. 2013. Nest and breeding population abundance of Least Terns: Assessing bias and variation in survey timing and methods. Journal of Field Ornithology 84: 287–298.

  • Hunt, K.L.*, D.H. Catlin, J.H. Felio* and, J.D. Fraser. 2013. The effect of capture frequency on the survival of Piping Plover chicks. Journal of Field Ornithology 84: 299–303. 

  • Hunt, K.L.*, N. Taygan, D.H. Catlin, J.H. Felio*, and J.D. Fraser. 2013. Demography of Snowy Plovers (Charadrius nivosus) on the Missouri River. Waterbirds 36(2): 220–224.

  • Catlin, D.H., J. H. Felio, and J.D. Fraser. 2013. Effects of water discharge on fledging time, growth, and survival of piping plovers on the Missouri River. Journal of Wildlife Management 77: 525–533.


  • Catlin, D.H., J.H. Felio*, and J.D. Fraser.  2012.  Comparison of piping plover foraging habitat on artificial and natural sandbars on the Missouri River.  Prairie Naturalist 44: 3–9.

  • Gratto-Trevor, C., D. Amirault-Langlais, D. Catlin, F. Cuthbert, J. Fraser, S. Maddock, E. Roche, and F. Shaffer. 2012. Connectivity in piping plovers: Do breeding populations have distinct winter distributions? Journal of Wildlife Management 76: 348–355.


  • Catlin, D.H., J.H. Felio*, and J.D. Fraser. 2011. Effect of great horned owl trapping on chick survival in piping plovers. Journal of Wildlife Management 75: 458-462.

  • Catlin, D.H.*, J.D. Fraser, J.H. Felio*, and J.B. Cohen. 2011. Piping plover habitat selection and nest success on natural, managed, and engineered sandbars. Journal of Wildlife Management 75: 305–310.

  • Cohen, J.B., B.D. Gerber, S.M. Karpanty, J.D. Fraser, and B.R. Truitt. 2011. Day and night foraging of Red Knots (Calidris canutus) during spring stopover in Virginia, USA. Waterbirds 34: 352-356.

  • Karpanty, S.M., J.B. Cohen, J. D. Fraser, and J. Berkson. 2011. Evidence of sufficient horseshoe crab eggs for a reduced red knot population during spring migration stopover in Delaware Bay USA. Journal of Wildlife Management 75: 984-994.


  • Catlin, D.H., R.B. Jacobson, M.H. Sherfy, M.J. Anteau, J.H. Felio, J.D. Fraser, C. Lott, T.L. Shaffer, and J.H. Stucker. 2010. Discussion of "Natural hydrograph of the Missouri River near Sioux City and the least tern and piping plover" by Donald Jorgensen. Journal of Hydrological Engineering 15: 1076–1078.

  • Cohen, J.B.*, J.D. Fraser. 2010. Piping Plover foraging distribution and prey abundance in the pre-laying period. Wilson Journal of Ornithology. 122: 578-582.

  • Cohen, J.B., S.M. Karpanty, and J.D. Fraser. 2010. Habitat selection and behavior of Red Knots (Calidris canutus) on the New Jersey Atlantic Coast during spring stopover. Condor 112: 665-662.

  • Fraser, J.D, S.M. Karpanty, and J.B. Cohen. 2010. Shorebirds forage disproportionately in horseshoe crab nest depressions. Waterbirds 33: 96-100.

  • Roche, E.A., J.B. Cohen, D.H. Catlin, D.L. Amirault-Langlais, F.J. Cuthbert, C.L. Gratto-Trevor, J. Felio*, and J.D. Fraser. 2010. Range-Wide Piping Plover Survival: Correlated Patterns and Temporal Declines. Journal of Wildlife Management 74: 1784–1791.

  • Cohen, J.B.*, L.M. Houghton, and J.D. Fraser. 2009. Nesting density and reproductive success of piping plovers in response to storm and human-created habitat changes. Wildlife Monographs. 173: 1-24


  • Cohen J.B. S.M. Karpanty, J.D. Fraser, B.D. Watts, and B.R. Truitt. 2009. Red knot stopover in Virginia, Delaware, and New Jersey: numbers, residency, and interchange. Journal of Wildlife Management 73: 939-945.

  • Cohen, J.B.*, S.M. Karpanty, D.H. Catlin*, J.D. Fraser, and R.A. Fischer. 2008. Winter ecology of piping plovers at Oregon Inlet, North Carolina. Waterbirds 31: 472–479.

  • Cohen, J.B.*, E.C. Wunker, and J.D. Fraser. 2008. Substrate and Vegetation selection by nesting Piping Plovers (Charadrius melodus) in New York. Wilson Journal of Ornithology 120:404-407.

  • Le Fer, D.*, J.D. Fraser, and C.D. Kruse. 2008. Piping plover chick foraging, growth, and survival in the Great Plains. Journal of Wildlife Management 72: 682-687.

  • Le Fer, D.*, J.D. Fraser, and C.D. Kruse. 2008. Piping plover foraging-site selection on the Missouri River. Waterbirds 31: 587-592.

  • Karpanty, S.M, J.D. Fraser, J. Berkson, L.J. Niles, A. Dey, and E. P. Smith. 2006. Horseshoe crab eggs determine red knot distribution in Delaware Bay habitats. Journal of Wildlife Management. 70: 1704-1710.

  • Cohen, J. B.*, J. D. Fraser, and D. H. Catlin*. 2006. Survival and site fidelity of Piping Plovers on Long Island, New York. Journal of Field Ornithology, 77(4): 409–417.

  • Fraser, J.D., S.H. Keane,* and P. A. Buckley. 2005. Prenesting use of intertidal habitats by piping plovers on South Monomoy Island. Journal of Wildlife Management 69: 1731-1736.

  • Elias, S.P.*, J.D. Fraser and P.A. Buckley. 2000. Piping plover brood foraging ecology on New York barrier Islands. Journal of Wildlife Management 64: 346-354.

  • Loegering, J.P.* and J.D. Fraser. 1995. Ghost crab preys on piping plover chick. Wilson Bulletin 107: 768-769.

  • Loegering, J.P.* and J.D. Fraser. 1995. Piping plover survival in different brood-rearing habitats. Journal of Wildlife Management 59: 646-655.

  • Patterson, M.E.*, J.D. Fraser, and J.W. Roggenbuck. Factors affecting piping plover productivity on Assateague Island. 1991. Journal of Wildlife Management 55: 526-531.

  • Patterson, M.E.*, J.D. Fraser, and J.W. Roggenbuck. 1990. Piping plover ecology, management and research needs. Virginia Journal of Science 41: 419-426.

*student author